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قراءة كتاب On the Relations of Man to the Lower Animals

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On the Relations of Man to the Lower Animals

On the Relations of Man to the Lower Animals

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دار النشر: Project Gutenberg
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being one of the yet unsolved problems of embryology), but the yelk becomes circumferentially indented, as if an invisible knife had been drawn round it, and thus appears divided into two hemispheres (Fig. 12, C).

By the repetition of this process in various planes, these hemispheres become subdivided, so that four segments are produced (D); and these, in like manner, divide and subdivide again, until the whole yelk is converted into a mass of granules, each of which consists of a minute spheroid of yelk-substance, inclosing a central particle, the so-called 'nucleus' (F). Nature, by this process, has attained much the same result as that at which a human artificer arrives by his operations in a brickfield. She takes the rough plastic material of the yelk and breaks it up into well-shaped tolerably even-sized masses, handy for building up into any part of the living edifice.

Fig. 14.--earliest Rudiment of the Dog. B. Rudiment Further Advanced, Showing the Foundations of The Head, Tail, And Vertebral Column. C. The Very Young Puppy, With Attached Ends of The Yelk-sac and Allantois, And Invested in the Amnion.

Next, the mass of organic bricks, or 'cells' as they are technically called, thus formed, acquires an orderly arrangement, becoming converted into a hollow spheroid with double walls. Then, upon one side of this spheroid, appears a thickening, and, by and bye, in the centre of the area of thickening, a straight shallow groove (Fig. 13, A) marks the central line of the edifice which is to be raised, or, in other words, indicates the position of the middle line of the body of the future dog. The substance bounding the groove on each side next rises up into a fold, the rudiment of the side wall of that long cavity, which will eventually lodge the spinal marrow and the brain; and in the floor of this chamber appears a solid cellular cord, the so-called 'notochord.' One end of the inclosed cavity dilates to form the head (Fig. 13, B), the other remains narrow, and eventually becomes the tail; the side walls of the body are fashioned out of the downward continuation of the walls of the groove; and from them, by and bye, grow out little buds which, by degrees, assume the shape of limbs. Watching the fashioning process stage by stage, one is forcibly reminded of the modeller in clay. Every part, every organ, is at first, as it were, pinched up rudely, and sketched out in the rough; then shaped more accurately; and only, at last, receives the touches which stamp its final character.

Thus, at length, the young puppy assumes such a form as is shown in Fig. 13, C. In this condition it has a disproportionately large head, as dissimilar to that of a dog as the bud-like limbs are unlike his legs.

The remains of the yelk, which have not yet been applied to the nutrition and growth of the young animal, are contained in a sac attached to the rudimentary intestine, and termed the yelk sac, or 'umbilical vesicle.' Two membranous bags, intended to subserve respectively the protection and nutrition of the young creature, have been developed from the skin and from the under and hinder surface of the body; the former, the so-called 'amnion,' is a sac filled with fluid, which invests the whole body of the embryo, and plays the part of a sort of water-bed for it; the other, termed the 'allantois,' grows out, loaded with blood-vessels, from the ventral region, and eventually applying itself to the walls of the cavity, in which the developing organism is contained, enables these vessels to become the channel by which the stream of nutriment, required to supply the wants of the offspring, is furnished to it by the parent.

The structure which is developed by the interlacement of the vessels of the offspring with those of the parent, and by means of which the former is enabled to receive nourishment and to get rid of effete matters, is termed the 'Placenta.'

It would be tedious, and it is unnecessary for my present purpose, to trace the process of development further; suffice it to say, that, by a long and gradual series of changes, the rudiment here depicted and described becomes a puppy, is born, and then, by still slower and less perceptible steps, passes into the adult Dog.

There is not much apparent resemblance between a barndoor Fowl and the Dog who protects the farm-yard. Nevertheless the student of development finds, not only that the chick commences its existence as an egg, primarily identical, in all essential respects, with that of the Dog, but that the yelk of this egg undergoes division—that the primitive groove arises, and that the contiguous parts of the germ are fashioned, by precisely similar methods, into a young chick, which, at one stage of its existence, is so like the nascent Dog, that ordinary inspection would hardly distinguish the two.

The history of the development of any other vertebrate animal, Lizard, Snake, Frog, or Fish, tells the same story. There is always, to begin with, an egg having the same essential structure as that of the Dog:—the yelk of that egg always undergoes division, or 'segmentation' as it is often called: the ultimate products of that segmentation constitute the building materials for the body of the young animal; and this is built up round a primitive groove, in the floor of which a notochord is developed. Furthermore, there is a period in which the young of all these animals resemble one another, not merely in outward form, but in all essentials of structure, so closely, that the differences between them are inconsiderable, while, in their subsequent course, they diverge more and more widely from one another. And it is a general law, that, the more closely any animals resemble one another in adult structure, the longer and the more intimately do their embryos resemble one another: so that, for example, the embryos of a Snake and of a Lizard remain like one another longer than do those of a Snake and of a Bird; and the embryo of a Dog and of a Cat remain like one another for a far longer period than do those of a Dog and a Bird; or of a Dog and an Opossum; or even than those of a Dog and a Monkey.

Thus the study of development affords a clear test of closeness of structural affinity, and one turns with impatience to inquire what results are yielded by the study of the development of Man. Is he something apart? Does he originate in a totally different way from Dog, Bird, Frog, and Fish, thus justifying those who assert him to have no place in nature and no real affinity with the lower world of animal life? Or does he originate in a similar germ, pass through the same slow and gradually progressive modifications,—depend on the same contrivances for protection and nutrition, and finally enter the world by the help of the same mechanism? The reply is not doubtful for a moment, and has not been doubtful any time these thirty years. Without question, the mode of origin and the early stages of the development of man are identical with those of the animals immediately below him in the scale:—without a doubt, in these respects, he is far nearer the Apes, than the Apes are to the Dog.

The Human ovum is about 1/125 of an inch in diameter, and might be described in the same terms as that of the Dog, so that I need only refer to the figure illustrative (14 A) of its structure. It leaves the organ in which it is formed in a similar fashion and enters the organic chamber prepared for its reception in the same way, the conditions of its development being in all respects the same. It has not yet been possible (and only by some rare chance can it ever be possible) to study the human ovum in so early a developmental stage as that of yelk division, but there is every reason to conclude that the changes it undergoes are identical with those exhibited by the ova of other vertebrated animals; for the formative materials of which the rudimentary

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