by the study of a small district. This should, if possible, be an island. If we took a slice of the continent like France or Germany, we should find the problem more complex. Instead of choosing the British Islands, we might, however, take an island like Corsica or Sardinia. In either of these we should discover peculiarities in the composition of their fauna precisely similar to those which I have indicated to be present in the British fauna. We should find probably a more striking endemic[3] element, which with us is so meagre that it can almost be left unnoticed; the main features, however, remain nearly the same. The fauna of both of these islands is composed of a strong southern element, of an eastern and a northern one, and in addition we have here species whose ancestors lived in Western Europe.

Before investigating more minutely the problems suggested by the composition of the faunas of these insular and also of some continental areas, it is necessary that we should thoroughly understand all about the migrations of animals. One of the principal objects of this work is to show how the autochthonous animals of Europe, i.e., those which have originated there, may be distinguished from the immigrants, and to trace the latter to the home of their ancestors. But in doing so, it is necessary to refer to the many important geographical changes which have occurred in Europe during the latest geological epochs. The study of the geographical distribution of the European fauna, as expounded in this work, will in many instances confirm the theories as to geographical changes based upon geological foundations. But in every case the views herein advocated are founded upon the geographical distribution of living and extinct organisms alone.

A terrestrial mammal like the deer can, under ordinary circumstances, only reach one part of a country from another by walking or running to it; but a beetle, such as the cockchafer, has two different modes of progression. It may walk or fly. In both, however, there is a third mode of transport—an involuntary one. The deer may be suddenly seized by a flood whilst crossing a river, and carried far away without necessarily coming to grief. The beetle in a similar manner could be transported to a distant country, or it might be caught in a whirlwind and blown hundreds of miles off.

We may thus distinguish between the natural or active and the accidental or passive means of distribution of animals. The active mode of dispersal again may be only migratory, as in most animals, or periodic and migratory, as in some birds and fishes. It is of course the tendency of every species to spread in all directions from its original home, provided it does not encounter obstacles, such as want of food, unsuitability of climate or soil, or barriers such as mountains, rivers, or the sea. Birds might be thought to be little interfered with by any of these barriers, but, as Dr. Wallace has shown, they are almost as much affected by them in their distribution as mammals are.

This then is the ordinary migratory distribution. Periodic distribution obtains with migratory birds and fishes. The annual flight of swallows to their northern summer residence comes under the heading of periodic migration or distribution, but apart from this, the swallow must seek to extend its range by the ordinary method, like every other animal. Similarly, the herring migrates periodically into shallow water to spawn, only to return again to its deeper home, where, as its numbers increase, there must be a tendency to spread. We have in these cases, therefore, both a periodic and an ordinary movement of migration.

Now, in studying the composition of a fauna, and especially its origin, it is of the utmost importance to be able to determine approximately the percentage of accidental arrivals and of the ordinary migrants—that is to say, of those which have reached the country owing to accidental distribution, and of animals which have adopted the usual course of migration. It is of all the more import to review this subject of accidental, or, as Darwin called it, "the occasional means" of distribution, as both he and Dr. Wallace have, I venture to think, somewhat over-estimated its significance. No one doubts that accidental transportal takes place, but the question is whether the accidentally transported animals arrive living and reach a spot where suitable food is procurable, and whether they are able to propagate their own species in the new locality. For it must be clear to anybody that the accidental transportal of a beetle or of a snail to a new country cannot affect its fauna or add one permanent member to it unless all these conditions are fulfilled. As a matter of fact, only exceedingly few instances are on record of man having witnessed, for example, the accidental transportal across the sea to an island of a live animal.

To mention an example, Colonel Feilden informs us (Zoologist, 1888) that, when living on the island of Barbadoes, an alligator arrived one day on the shore, and at the same time a tree measuring 40 feet in length, which was recognised as a Demerara species, was likewise stranded. He thinks that there can be no doubt that the alligator, which was alive when it reached Barbadoes, was transported by the tree, thus covering a distance of 250 miles from the nearest land. Numerous observations on the accidental transportal of seeds and tree-trunks from one island to another, and from a continent to an island, have been recorded, and even on our own shores we may witness the occasional arrival of such vegetable products from a far distant land. On the west coast of Ireland it not unfrequently happens that large West Indian beans are stranded, and in this as well as in many other similar cases the seeds have often proved none the worse for their prolonged immersion in sea-water. That locusts are sometimes blown to great distances from the land is not so surprising, since their power of steering through the air is very limited. Darwin mentions (p. 327) having caught one 370 miles from the coast of Africa, and that swarms of them sometimes visited Madeira. Sir Charles Lyell relates that green rafts composed of canes and brush-wood are occasionally carried down the Parana River in South America by inundations, bearing on them the tiger, cayman, squirrels, and other quadrupeds.

But though actual observations of such abnormal instances of the dispersal of animals are few, many experiments have been made to demonstrate the possibility of a passive transportal of species over wide distances. It was especially Darwin who gave a great stimulus by setting the example to those interested in natural history in the conduct of such researches. He was struck by the fact that, though land-shells and their eggs are easily killed by sea-water, almost all oceanic islands, even the smallest and most isolated, are inhabited by them, and felt that there must be some unknown but occasionally efficient means for their transportal (p. 353). To quote his words: "It occurred to me that land-shells, when hibernating and having a membranous diaphragm over the mouth of the shell, might be floated in chinks of drifted timber across moderately wide arms of the sea. And I find that several species in this state withstand uninjured an immersion in sea-water during seven days: one shell, the Helix pomatia, after having been thus treated and again hibernating, was put into sea-water for twenty days, and perfectly recovered. During this length of time the shell might have been carried by a marine current of average