قراءة كتاب Morphological Variation in a Population of the Snake, Tantilla gracilis Baird and Girard
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Morphological Variation in a Population of the Snake, Tantilla gracilis Baird and Girard
plus one-half.
The anterior and posterior edges of the suborbital process were determined as parallel or not at any point by aligning them with parallel lines on an ocular grid (Fig. 5b).
The angle of the posterior edge of the lateral flange was determined as an acute, obtuse, or right angle by fitting it to a pair of perpendicular lines of an ocular grid (Fig. 5b).
The number of lateral anterior foramina (Fig. 4b) was determined. If only two are present, their lengths (anterior to posterior) were compared by measuring them with an ocular micrometer. Also, if two lateral anterior foramina are present, the point of intersection of two perpendicular lines of an ocular grid determined which tooth-socket occurs directly below the center of the posterior foramen (Fig. 4b).
RESULTS
General Characteristics
From the data given by Force (1935), Taylor (1936), and Kirn et al. (1949), and from our own data, we conclude that a typical specimen of gracilis, meaning a specimen having the usual characteristics of the species, has the following characteristics in combination: supralabials 6-6; infralabials 6-6; preoculars 1-1; postoculars 1-1; temporals 1 + 1 on each side; nasal divided below naris; supralabials 3 + 4 entering orbit; mental in contact with chin-shields; all five dorsal scale counts 15; and plate divided. (In the present work we use the words usual, unusual, typical, and atypical with neither quotation marks nor apology).
Of the 244 specimens on which we could examine each of these characteristics, only 125 (51.2%) have all of the typical characteristics in combination. The 119 specimens that do not possess all of the typical characteristics in combination include 31 (12.7% of the 244) that lack at least two of the typical characteristics.
Sex
The sample (N = 246) is composed of 107 (43.5%) females and 139 (56.5%) males.
Measurements
Females have total lengths from 96 to 244 mm. (mean, 173.3; N = 79). One female (KU 83480) measuring 244 mm. may be the longest specimen known (Conant, 1958, reports the longest as 9-1/8 inches—approximately 232 mm.). Males have total lengths from 96 to 215 mm. (mean, 162.4; N = 109). Females have tail lengths from 16 to 50 mm. (mean, 34.6; N = 79), and males have tail lengths from 21 to 53 mm. (mean, 37.8; N = 109).
The ratio of tail length to total length in females is from 0.17 to 0.22 (mean, 0.20; N = 79); in males it is from 0.21 to 0.27 (mean, 0.23; N = 109; Fig. 1). Juveniles (those less than 125 mm. in total length according to Force, 1935: tables 1 and 2) have proportionally shorter tails than do adults; the ratio of tail length divided by total length is 0.17-0.21 (mean, 0.18; N = 14) in females and 0.21-0.23 (mean, 0.22; N = 16) in males. Adult females (125 mm. or more in total length) have ratios from 0.18 to 0.22 (mean, 0.20; N = 65) and adult males have ratios from 0.21 to 0.27 (mean, 0.23; N = 93).
Of the 246 specimens examined, 58 have incomplete tails. Of these 58 specimens, six had freshly broken tails, so we assume that 52 (21.1% of the 246) have incomplete tails resulting from natural causes; the remaining six specimens
could have had their tails broken at or soon after capture, assuming that none of the snakes was retained in captivity for an extended period.
Supralabials
The supralabials are either six (usually) or seven on a side (Table 1). When seven supralabials are present on either one or both sides, the addition results from one of the following conditions: a) 5 is divided to form 5 and 6 (N = 6); b) 2 and 3 form 2, 3, and 4 (N = 1); c) 5 and 6 form 5, 6, and 7 (N = 12); d) 4, 5, and 6 form 4, 5, 6, and 7 (N = 1); e) 1 and 2 form 1, 2, and 3 (N = 1); f) 1, 2, and 3 form 1, 2, 3, and 4 (N = 1); g) 2 is fused with the nasal, and 3 and 4 are reduced to granules (N = 1); h) type of addition unknown (N = 5).
Table 1. Variation in Number of Supralabials (N = 245) and Infralabials (N = 246) of Tantilla gracilis.
Number of Supralabials |
Number of specimens |
Per cent of occurrence |
Number of infralabials |
Number of specimens |
Per cent of occurrence |
---|---|---|---|---|---|
6-6 | 221 | 90.2 | 5-5 | 5 | 2.0 |
6-7 | 10 | 4.1 | 5-6 | 4 | 1.6 |
7-6 | 10 | 4.1 | 6-5 | 5 | 2.0 |
7-7 | 4 | 1.6 | 6-6 | 229 | 93.1 |
6-7 | 1 | 0.4 | |||
7-6 | 2 | 0.8 |
In three of the four specimens with 7-7 supralabials, the type of addition is the same on each side, whereas the other specimen has a different type of addition on each side. Of the 23 examples for which the type of addition is known, 19 (82.6%) have the increase in number on the posterior half of the series.
In one specimen the third supralabial on the right is diagonally divided and the resulting upper part does not contact the edge of the lip; hence the specimen has 6-6 supralabials.
Infralabials
The infralabials are five, six (usually), or