long, branched hairs which cover this region become quite moist and sticky, since the brushes of these two pair of legs are wet and the pollen which they bear possesses a superabundance of the moistening fluid. Any dry pollen which passes over this region and touches these hairs receives moisture by contact with them. This is particularly true of the free dry pollen which the middle pair of legs collect by combing over the sides of the thorax.

Fig. 6.—A bee upon the wing, showing the position of the middle legs when they touch and pat down the pollen masses. A very slight amount of pollen reaches the corbiculæ through this movement. (Original.)

The pollen upon the middle legs is transferred to the hind legs in at least two ways. By far the larger amount is deposited upon the pollen combs which lie on the inner surfaces of the plantæ of the hind legs. To accomplish this a middle leg is placed between the plantæ of the two hind legs, which are brought together so as to grasp the brush of the middle leg, pressing it closely between them, but allowing it to be drawn toward the body between the pollen combs of the two hind legs. (See fig. 5.) This action results in the transference of the pollen from the middle-leg brush to the pollen combs of the hind leg of the opposite side, since the combs of that leg scrape over the pollen-laden brush of the middle leg. This action may take place while the bee is on the wing or before it leaves the flower.

The middle legs place a relatively small amount of pollen directly upon the pollen masses in the corbiculæ. This is accomplished when the brushes of the middle legs are used to pat down the pollen masses and to render them more compact. (See fig. 6.) The legs are used for this purpose quite often during the process of Loading the baskets, and a small amount of pollen is incidentally added to the masses when the brushes come into contact with them. A misinterpretation of this action has led some observers into the erroneous belief that all or nearly all of the corbicular pollen is scraped from the middle-leg brushes by the hairs which fringe the sides of the baskets. The middle legs do not scrape across the baskets, but merely pat downward upon the pollen which is there accumulating.

It is also possible that, in transferring pollen from the middle leg of one side to the planta of the opposite hind leg, the middle-leg brush may touch and rub over the pecten of the hind leg and thus directly place some of its pollen behind the pecten spines. Such a result is, however, very doubtful.

ACTION OF THE HIND LEGS.

The middle legs contribute the major portion of the pollen which reaches the hind legs, and all of it in cases where all of the pollen first reaches the bee in the region of the mouth. However, when much pollen falls upon the body of the bee the hind legs collect a little of it directly, for it falls upon their brushes and is collected upon them when these legs execute cleansing movements to remove it from the ventral surface and sides of the abdomen. All of the pollen which reaches the corbiculæ, with the exception of the small amount placed there by the middle legs when they pat down the pollen masses, passes first to the pollen combs of the plantæ.

When in the act of loading pollen from the plantar brushes to the corbiculæ the two hind legs hang beneath the abdomen with the tibio-femoral joints well