قراءة كتاب An Introduction to Entomology: Vol. III (of 4) or Elements of the Natural History of the Insects
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An Introduction to Entomology: Vol. III (of 4) or Elements of the Natural History of the Insects
great Aristotle of the moderns, Linné.
This illustrious naturalist, aware of the incorrectness of the primary divisions of the animal kingdom founded upon the presence or absence of blood, establishes his system upon the structure of the heart, and upon the temperature and colour of the circulating fluid. He divided animals into two great sections or sub-kingdoms, each comprising two classes. His first section included those having a heart with two ventricles, two auricles, and warm and red blood, viz. the Mammalia or beasts, and the Aves or birds. His second, those having a heart with one ventricle, one auricle, and cold and red blood, namely, the classes Amphibia, which included reptiles, serpents, &c. and Pisces or fish. His third, those having a heart with one ventricle and no auricle, and cold white sanies in the place of blood, namely, his classes Insecta et Vermes, including the Invertebrate animals of Lamarck. Thus the first of Aristotle's great divisions he increased by the addition of a new and very distinct class, the Amphibia, by which some ground was gained in the science; but as much was lost by his compressing the four classes of which the last consisted into two, by which the natural classes of Cephalopoda and Crustacea merged under Insecta and Vermes. Linné was not aware of the extraordinary fact, that the Cephalopoda have three hearts; and that though the Crustacea and Arachnida have a circulation, Insects have none, or he would never have taken this retrograde step.
Indeed Linné's definition of an Insect is, in many most material points, inapplicable, not only to the Crustacea, but to many other animals included under that denomination. This will appear evident from a very slight examination. Thus it runs: "Polypod animalcula, breathing by lateral spiracles, armed every where with an osseous skin, whose head is furnished with movable sensitive antennæ[10]." Now of this definition only the first member can be applied to the whole class which it is meant to designate; for the entire genus Cancer L., which, with some others, forms the class Crustacea of the moderns, does not respire by spiracles at all, but by gills; and the same in some degree may be said of spiders, scorpions, &c. With the last member of the definition Linné himself must have been aware that a large number of what he conceived to be insects were at variance, as mites, spiders, and many other of his apterous tribes: though from some very recent observations of M. Latreille[11], there seems some ground for thinking, that in these the antennæ are represented by the mandibles, palpi, &c.[12], and to the soft flexible, coriaceous or membranous skin of a vast number of insects, the term cutis ossea is by no means applicable.
Evident as these incongruities are, when the Herculean task which Linné imposed upon himself, and the vastness and variety of his labours, are considered, they become very venial. Indeed, unless he had divided his class Insecta into two or more, it was impossible to define it intelligibly to ordinary readers, otherwise than nearly in the terms which he actually employed; and these characters, restricted and amended by qualifying clauses, are still those to which recurrence must be had in a popular definition of the class, when separated as it ought to be from the Crustacea and Arachnida.
Pennant, Brisson, and other zoologists, who, attending to nature rather than system, saw the impropriety of uniting a crab or a lobster in the same class with a bee or a beetle, long since assigned the Crustacea their ancient distinct rank. "But these changes," as Latreille observes[13], "being only founded upon external characters, might be deemed arbitrary; and to fix our opinion, it was necessary to have recourse to a decisive authority—the internal and comparative organization of these animals." It results from the observations of the most profound comparative anatomist of our age, M. Cuvier, that the Crustacea and Arachnida differ from insects properly so called, and particularly from those that are furnished with wings, in having a complete system of circulation, a different mode of respiration, and that they have a more perfect organization. Influenced by these motives, both Cuvier and Lamarck have considered them as forming two classes separate from insects. Treviranus, led by considerations founded on the organs of circulation, of respiration, and of generation, is of opinion that spiders and scorpions ought to form one class with the Crustacea: he observes, however, that the nervous system of all three is very dissimilar; and that in an arrangement founded on this circumstance, the organs of motion, and the external shape, even spiders and scorpions must be placed in different classes[14].
It is to be observed with regard to the Arachnida of the French school, that the class as laid down by them includes several animals that have no circulation, and breathe by tracheæ, of which description are the mites (Acarus L.), and the harvest-men (Phalangium L.) &c.; and therefore it has been divided into two orders, Pulmonaria and Tracheana; but if the definition from the internal organization be adhered to, the latter should either remain with the class Insecta, or form a new one by themselves. Yet the animals that compose the Trachean order of Arachnida, their external form considered, are certainly much more nearly related to the spiders and scorpions than to any members of the class Insecta at present known. This circumstance, perhaps, may seem to throw some doubt upon the modern system of classification.
I must further observe, that the assertion of Treviranus, which appears to intimate that the respiration of the pulmonary Arachnida is the same with that of the Crustacea, is not quite correct, since in the latter the branchiæ or gills are external, and in the former internal, the air entering by spiracles before it acts upon them[15].
It may not be amiss in this place to lay before you the principal points in which the Crustacea and Arachnida agree with Insecta, and also those in which they differ.
The Crustacea agree with Insecta in having a body divided into segments, furnished with jointed legs, compound eyes, and antennæ. Their