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قراءة كتاب The Beaked Whales of the Family Ziphidae An account of the Beaked Whales of the Family Ziphiidae in the collection of the united states museum...
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The Beaked Whales of the Family Ziphidae An account of the Beaked Whales of the Family Ziphiidae in the collection of the united states museum...
bone occupies the bottom of the maxillary notch and a small portion of it is visible from above, while in bidens it does not extend up into the notch at all from the inferior surface and is not visible from above. The posterior margin of the maxillæ is more squared in europæus than in bidens.
The margins of the beak, formed by the maxillæ, instead of being straight, are somewhat emarginate a little posterior to the middle of the length and somewhat convex anterior to it, which gives the contour of the beak, seen from above, a different shape from that of bidens. In the type-skull of europæus the mesirostral ossification appears to be higher at the proximal end than the premaxillæ, and distally extends to the end of the beak. In bidens it is lower than the premaxillæ and, in the Nantucket skull at least, ends anteriorly at the same point as the vomer, or, in other words, much behind the end of the beak. It would appear from the statements of Sir William Turner, Van Beneden and Gervais, Grieg, and others, that the mesirostral ossification never reaches the end of the beak in bidens, but it does in grayi, haasti, densirostris, and many fossil species, as well as in europæus.
Lateral aspect (Pl. 8, figs. 1, 2).—The temporal fossæ are a little longer than the orbit in europæus, but a little shorter than the orbit in bidens; in the former the superior margin is flat or a little concave, rather than convex. The exoccipital extends in an angle farther forward in europæus, and the suture between it and the zygomatic is, in consequence, less nearly vertical than in bidens. The premaxillæ at the sides of the blowhole are nearly horizontal, so that their superior surface is little seen from this aspect, while in bidens they slope downward, so that the whole of the superior surface is visible. The high maxillary ridge, situated behind the anteorbital notch, is very noticeable from this point of view, as it shuts off a considerable portion of the premaxillæ. The convex inferior outline of the beak and its great depth at the base are also salient peculiarities.
Ventral aspect (Pl. 5, figs. 1, 2).—The anterior ends of the palatine bones are bifurcated, the inner part being the smaller. The two bones make but a narrow angle with the median line, instead of a wide one, as in bidens, and the surface of the maxillæ between them is strongly convex instead of flat. This convexity is narrowed at both ends, or, in other words, is fusiform in shape. No similar conformation is found in bidens, in which the inferior basal area of the maxillæ is flat.
In the young Atlantic City skull of europæus, the vomer is visible as a small, narrow, club-shaped piece, 68 mm. long. Anteriorly it joins the premaxillæ, which form a prominent ridge in the median line. On each side of this ridge is a wide and quite deep groove. As the beak is lacking in the adult North Long Branch skull, its peculiarities can not be made known. In the type-skull the form is the same as in the Atlantic City skull, but the vomer does not appear at all on the palate. In bidens the shape of the inferior surface of the premaxillæ at the distal end is quite different. A very narrow groove runs parallel with and close to the median line and the whole surface external to it is more or less convex.
MANDIBLE.
The mandible of the Atlantic City specimen of M. europæus resembles that of the type, as figured by Van Beneden and Gervais, in the shortness of the symphysis and in the position of the tooth, which is in advance of the posterior end of the symphysis. A number of differences, however, require consideration. (Pl. 11, figs. 3 and 6.)
In the type, the symphysis, as shown by Van Beneden and Gervais’ figure, plate 24, fig. 2a, is a little more than one-fifth the length of the mandible. The same relative proportion is found in the Atlantic City specimen, but, as the latter is a younger individual, one would expect the symphysis to be shorter. The figure of Van Beneden and Gervais gives the impression that in the type the end of the mandible is broken, and that, hence, the symphysis is shorter than it was originally. It will be observed that figures 2 and 2a do not agree as regards the length between the tooth and the end of the jaw, figure 2a showing a greater length. In figure 2, however, the jaw seems rather too long for the cranium, and if the greater length of the symphysis shown in figure 2a were introduced, it would certainly be so. The explanation of this discrepancy is not readily found; but one may be allowed to think that the symphysis is not so blunt in the type as is shown in figure 2.
In the Atlantic City specimen the superior lateral free margin of the symphysis is straight, while in the type it is much elevated. This is no doubt due to difference in age and possibly in sex. The type shows three or four mental foramina, while the Atlantic City specimen has one large posterior one and seven smaller ones anterior to it.
Another peculiarity of the latter specimen is that the coronoid process is situated much in advance of the condyle, while the angle extends considerably behind it. In the type both are nearly in line with the condyle. I am unable to explain this difference.
In the Atlantic City specimen the axis of the tooth where it emerges from the alveolus is 91 mm. from the end of the jaw. The portion of the tooth above the alveolus is 11 mm. long at the base and 12 mm. high. It is conical and sharp pointed, and is inclined forward and a little outward, especially at the tip. At the alveolus the transverse breadth of the tooth is 5 mm. The much larger tooth in the type indicates that that specimen was a male.
The mandible of the Atlantic City specimen of M. europæus differs from that of M. bidens in the relative shortness of the symphysis, the large number of mental foramina, the more anterior position of the tooth, and the direction of the crown, which is forward instead of backward.
Dimensions of the type and two other skulls of Mesoplodon europæus.
- Column headings:
- A: English Channel, type,a adult.
- B: North Long Branch, New Jersey, female, adult.
- C: Atlantic City, New Jersey, 23346 U.S.N.M., male, young.
Measurements. | A | B | C |
---|---|---|---|
mm. | mm. | mm. | |
Total length | 762 | (b) | 675 |
Length of rostrum | 459 | ... | 427 |
Tip of beak to posterior end of pterygoids | 561 | ... | 525 |
Height from vertex to end of pterygoids | c292? | 283 | 256 |
Breadth between orbits | 327 | d325 | d287 |
Breadth between zygomatic processes | 360 | e325 |