vertical, the interior has become filled up by a coat of sandstone, whilst the bark has become transformed into an envelope of an inch, or half an inch of coal. But many are found lying in the strata in a horizontal plane. These have been cast down and covered up by an ever-increasing load of strata, so that the weight has, in the course of time, compressed the tree into simply the thickness of the double bark, that is, of the two opposite sides of the envelope which covered it when living.

Sigillarae grew to a very great height without branching, some specimens having measured from 60 to 70 feet long. In accordance with their outside markings, certain types are known as syringodendron, favularia, and clathraria. Diploxylon is a term applied to an interior stem referable to this family.

[Illustration: FIG. 16.—Stigmaria ficoides. Coal-shale.]

But the most interesting point about the sigillariae is the root. This was for a long time regarded as an entirely distinct individual, and the older geologists explained it in their writings as a species of succulent aquatic plant, giving it the name of stigmaria. They realized the fact that it was almost universally found in those beds which occur immediately beneath the coal seams, but for a long time it did not strike them that it might possibly be the root of a tree. In an old edition of Lyell's "Elements of Geology," utterly unlike existing editions in quality, quantity, or comprehensiveness, after describing it as an extinct species of water-plant, the author hazarded the conjecture that it might ultimately be found to have a connection with some other well-known plant or tree. It was noticed that above the coal, in the roof, stigmariae were absent, and that the stems of trees which occurred there, had become flattened by the weight of the overlying strata. The stigmariae on the other hand, abounded in the underclay, as it is called, and were not in any way compressed but retained what appeared to be their natural shape and position. Hence to explain their appearance, it was thought that they were water-plants, ramifying the mud in every direction, and finally becoming overwhelmed and covered by the mud itself. On botanical grounds, Brongniart and Lyell conjectured that they formed the roots of other trees, and this became the more apparent as it came to be acknowledged that the underclays were really ancient soils. All doubt was, however, finally dispelled by the discovery by Mr Binney, of a sigillaria and a stigmaria in actual connection with each other, in the Lancashire coal-field.

Stigmariae have since been found in the Cape Breton coal-field, attached to Lepidodendra, about which we have already spoken, and a similar discovery has since been made in the British coal-fields. This, therefore, would seem to shew the affinity of the sigillaria to the lepidodendron, and through it to the living lycopods, or club-mosses.

Some few species of stigmarian roots had been discovered, and various specific names had been given to them before their actual nature was made out. What for some time were thought to be long cylindrical leaves, have now been found to be simply rootlets, and in specimens where these have been removed, the surface of the stigmaria has been noticed to be covered with large numbers of protuberant tubercles, which have formed the bases of the rootlets. There appears to have also been some special kind of arrangement in their growth, since, unlike the roots of most living plants, the tubercles to which these rootlets were attached, were arranged spirally around the main root. Each of these tubercles was pitted in the centre, and into these the almost pointed ends of the rootlets fitted, as by a ball and socket joint.

[Illustration: FIG. 17—Section of stigmaria.]

"A single trunk of sigillaria in an erect forest presents an epitome of a coal-seam. Its roots represent the stigmaria underclay; its bark the compact coal; its woody axis, the mineral charcoal; its fallen leaves and fruits, with remains of herbaceous plants growing in its shade, mixed with a little earthy matter, the layers of coarse coal. The condition of the durable outer bark of erect trees, concurs with the chemical theory of coal, in showing the especial suitableness of this kind of tissue for the production of the purer compact coals."—(Dawson, "Structures in Coal.")

There is yet one other family of plants which must be mentioned, and which forms a very important portion of the constituent flora of the coal period. This is the great family of the coniferae, which although differing in many respects from the highly organised dicotyledons of the present day, yet resembled them in some respects, especially in the formation of an annual ring of woody growth.

The conifers are those trees which, as the name would imply, bear their fruit in the form of cones, such as the fir, larch, cedar, and others. The order is one which is familiar to all, not only on account of the cones they bear, and their sheddings, which in the autumn strew the ground with a soft carpet of long needle-like leaves, but also because of the gum-like secretion of resin which is contained in their tissues. Only a few species have been found in the coal-beds, and these, on examination under the microscope, have been discovered to be closely related to the araucarian division of pines, rather than to any of our common firs. The living species of this tree is a native of Norfolk Island, in the Pacific, and here it attains a height of 200 feet, with a girth of 30 feet. From the peculiar arrangement of the ducts in the elongated cellular tissue of the tree, as seen under the microscope, the fossil conifers, which exhibit this structure, have been placed in the same division.

The familiar fossil known to geologists as Sternbergia has now been shown to be the cast of the central pith of these conifers, amongst which may be mentioned cordaites, araucarites, and dadoxylon.. The central cores had become replaced with inorganic matter after the pith had shrunk and left the space empty. This shrinkage of the pith is a process which takes place in many plants even when living, and instances will at once occur, in which the stems of various species of shrubs when broken open exhibit the remains of the shrunken pith, in the shape of thin discs across the interval cavity.

We might reasonably expect that where we find the remains of fossil coniferous trees, we should also meet with the cones or fruit which they bear. And such is the case. In some coal-districts fossil fruits, named cardiocarpum and trigonocarpum, have been found in great quantities, and these have now been decided by botanists to be the fruits of certain conifers, allied, not to those which bear hard cones, but to those which bear solitary fleshy fruits. Sir Charles Lyell referred them to a Chinese genus of the yew tribe called salisburia. Dawson states that they are very similar to both taxus and salisburia.. They are abundant in some coal-measures, and are contained, not only in the coal itself, but also in the sandstones and shales. The under-clays appear to be devoid of them, and this is, of course, exactly what might have been expected, since the seeds would remain upon the soil until covered up by vegetable matter, but would never form part of the clay soil itself.

In connection with the varieties which have been distinguished in the families of the conifers, calamites, and sigillariae, Sir William Dawson makes the following observations: "I believe that there was a considerably wide range of organisation in cordaitinae as well as in calamites and sigillariae, and that it will eventually be found that there were three lines of connection between the higher cryptogams