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   Failure of Female No. 11 to Whelp 19    Demise of the Translocated Wolves 19 DISCUSSION 21    Effect of Captivity and Human Contact 21    Movements 22       Environmental Influences 22       Possible Homing Tendencies 22       Distances Traveled 23       Home Range Size 25       Selection of a Territory 25    Vulnerability and Mortality 25    Food Habits and Predation 26    An Alternate Approach 26 CONCLUSIONS 26 ACKNOWLEDGMENTS 27 LITERATURE CITED 27

ABSTRACT

Two male and two female eastern timber wolves (Canis lupus lycaon), live-trapped in Minnesota were released in March 1974 near Huron Mountain in the Upper Peninsula of Michigan. Their movements were monitored by aerial radio-telemetry.

The wolves separated into a group of three and a single animal after release. The single, a young female, remained in the release region in an area of 346 square miles (896 km²). The pack of three moved generally westward for 13 days and then explored a 1,631 square-mile (4,224 km²) region but settled after 2 months in a 246 square-mile (637 km²) area about 55 miles (88 km) southwest of the release site. The adult female, which mated while captive prior to release, failed to whelp.

In early July, one male was killed by an automobile, and the other was shot. The remaining female from the pack then began to move over a much larger area again. On September 20th she was trapped by a coyote (Canis latrans) trapper and shot. Two months later the single female was killed by a deer (Odocoileus virginianus) hunter.

These results indicated that wolves can be transplanted to a new region, although they may not settle in the release area itself. The displacement of the translocated wolves in this experiment apparently caused an initial increase in their daily movements, and probably increased their vulnerability, at least during the first 2 months after release. The two females examined post-mortem were in good physical condition indicating that food supplies were adequate in Michigan.

Human-caused mortality was responsible for the failure of the wolves to establish themselves. Therefore recommendations for a more successful re-establishment effort include a stronger public-education campaign, removal of the coyote bounty, and release of a greater number of wolves.

INTRODUCTION

The eastern timber wolf (Canis lupus lycaon) originally occurred throughout the eastern United States and Canada but is now extinct in most of the United States. The only substantial population left inhabits northern Minnesota (Fig. 1). The estimated wolf population in the Superior National Forest of northeastern Minnesota in winter 1972–73 was about 390 (Mech 1973), and a tentative population estimate for the entire state is 500 to 1,000 (Mech and Rausch 1975). A well known population of about 15 to 30 wolves is also found in Isle Royale National Park, Lake Superior, Michigan (Mech 1966; Wolfe and Allen 1973; Peterson 1974).

In the Upper Peninsula of Michigan, Hendrickson et al. (1975) estimated the wolf population in 1973 at 6 to 10 animals, existing in three scattered areas: Iron County, Northern Marquette County, and Chippewa and Mackinac Counties (Fig. 2). Lone wolves made up 90 per cent of verified wolf observations there in recent years, and no more than two animals have been found together in at least the past 13 years.

Hendrickson et al. (1975) postulated that the current low wolf population is maintained through possible sporadic breeding and immigration from Ontario and Minnesota (via